Mitochondria directly interact with the nuclear pore complex; VDAC1 was the top mitochondrial candidate that interacts with the filamentous nuclear pore protein RANBP2.
www.nature.com/articles/s41...
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Mitochondria directly interact with the nuclear pore complex - Nature
Mitochondria interact directly with the nuclear pore complex via VDAC1–RANBP2 binding to sustain nuclear ATP levels.